Serpentes (snakes)
Serpentes - snakes
Lepidosauria; Squamata; Serpentes |
The Serpentes,
commonly known as snakes,
are a familiar and well defined group, whose monophyly has
strong support from a suite of both morphological and molecular characteristics
(see below). Their position within the squamates, however, has proved extremely
difficult to resolve. For more information regarding higher level squamate phylogeny, pleas refer to the Squamata main page.
Diversity and Lower Taxonomy:
Extant snakes
are divided into two well-supported monophyletic sister
groups: the Scolecophidia and the Alethinophidia.
Description:
Scolephidians are a poorly
known group of snakes that are small, fossorial, and worm-like.
Alethinophidians, however, are the more familiar group, possessing what
would generally be considered a snake-like body. They are generally larger and
less fossorial.
Features:
- Limbless, but many retain
traces of a pectoral girdle. Members
of some of the more primitive families, such as the boas and pythons (contained
within the superfamily Booidea), show external traces in the form of vestigial hindlimbs, called anal spurs, which flank the cloacal opening and are now
only used for clasping during courtship.
- Well developed chemosensation, with forked
tongue.
- Highly kinetic skull with eight
points of rotation, allowing large prey to be swallowed whole. Each side
of the skull can move independently.
- Many elements of the skull
reduced or lost, facilitating the evolution of kinesis.
- The two sides of the mandible are loosely
connected with cartilage at the rostral midline (in other
jawed vertebrates, the two sides of the mandible are strongly fused to
form the mandibular symphysis).
- Recurved teeth, preventing the
escape of seized prey victims.
- No external ear openings.
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